TYPE: Costa Rica. Cartago: Casamata, Tejar de San Carlos, collected by A. Flores R., 3 Feb 1991, flowered in cultivation at Jardín Botánico Lankester, Paraíso de Cartago, 1400 m, 17 May 1999, F. Pupulin 1463 (holotype, USJ; isotype, SEL; illustration of type).